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Rubia cordifolia var. pratensis inhabits sandy
soil on the fringe of low land forests close to the seashore. It
possesses 5-8 verticillate leaves around the main stem. In the
species level, that is, with Rubia cordifolia, leaves are
verticillate by four. The petioles, leaf margins, and veins on a
leaf bottom have lots of thick stickles.
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